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Post 24670

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Practically when naturalists are at work, they donot trouble themselves about the physiological value of the characters which they use in defining a group, or in allocating any particular species. If they find a character nearly uniform, and common to a great number of forms, and not common to others, they use it as one of high value; if common to some lesser number, they use it as of subordinate value. This principle has been broadly confessed by some naturalists to be the true one; and by none more clearly than by that excellent botanist, Aug. St. Hilaire. If certain characters are always found correlated with others, though no apparent bond of connexion can be discovered between them, especial value is set on them. As in most groups of animals, important organs, such as those for propelling the blood, or for aƫrating it, or those for propagating the race, are found nearly uniform, they are considered as highly serviceable in classification; but in some groups of animals all these, the most important vital organs, are found to offer characters of quite subordinate value.

We can see why characters derived from the embryo should be of equal importance with those derived from the adult, for our classifications of course include all ages of each species. But it is by no means obvious, on the ordinary view, why the structure of the embryo should be more important for this purpose than that of the adult, which alone plays its full part in the economy of nature. Yet it has been strongly urged by those great naturalists, Milne Edwards and Agassiz, that embryonic characters are the most important of any in the classification of animals; and this doctrine has very generally been admitted as true. The same fact holds good with flowering plants, of which the two main divisions have been founded on characters derived from the embryo,–on the number and position of the embryonic leaves or cotyledons, and on the mode of development of the plumule and radicle. In our discussion on embryology, we shall see why such characters are so valuable, on the view of classification tacitly including the idea of descent.

Our classifications are often plainly influenced by chains of affinities. Nothing can be easier than to define a number of characters common to all birds; but in the case of crustaceans, such definition has hitherto been found impossible. There are crustaceans at the opposite ends of the series, which have hardly a character in common; yet the species at both ends, from being plainly allied to others, and these to others, and so onwards, can be recognised as unequivocally belonging to this, and to no other class of the Articulata.

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